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During the ISR, with global protein translation suppressed, untranslated mRNA transcripts and aggregation-prone RNA binding proteins such as Ras GTPase-activating protein-binding protein 1/2 (G3BP1/2) and T-cell-restricted intracellular antigen-1 related protein (TIAR) are packaged into non-membrane bound foci known as stress granules (SGs) [8]

J Biol Chem 276: 2099921003

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Its defining modification is a trans -3-hexenoyl group attached to the N-terminal tyrosine, a chemical change designed to protect the peptide from rapid cleavage by dipeptidyl peptidase-4 (DPP-4), the enzyme that quickly inactivates native GHRH